Department of Forest Sciences, Colorado State University, Fort Collins, CO.    1988 1992

    Ph.D. in Forest Ecology.

 

  School of Natural Resources and the Environment, University of Michigan, Ann Arbor, MI. 1986 1988

    M.S. in Natural Resources.

 

  Department of Forestry, Nanjing Forestry University, Nanjing, P. R. China.1978 1982

    B. A. in Forest Science.

 

 

 

Ecosystem and soil ecology, especially ecological rhythms and interactions of soil organisms and ecosystem processes.

I am also interested in the transport and transformation of carbon along the land-river-ocean continuum.

 

 
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Ruan, H. H., X. M. Zou, F. N. Scatena, and J. K. Zimmerman. In press. Asynchronous rhythms of soil microbial biomass and plant litterfall in a tropical wet forest. Plant and Soil.

 

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Grau H. R., T. M. Aide, J. K. Zimmerman, J. R. Thomlinson, E. Helmer, X. M. Zou. 2003. The ecological consequences of socioeconomic and land use changes in post agriculture Puerto Rico. BioScience 53: 1159-1168.
 

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Sanchez, Y., X. M. Zou, S. Borges, and H. H. Ruan. 2003. Recovery of native earthworms in abandoned tropical pastures. Conservation Biology 17: 999-1006.
 

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Warren, M., and X. M. Zou. 2002. Tree species effect on soil macrofauna in tropical tree plantations of Puerto Rico. Forest Ecology and Management 170: 161-171.
 

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Liu, Z.G. and X. M. Zou. 2002. Exotic earthworms accelerate plant litter decomposition in a Puerto Rican pasture and a wet forest. Ecological Applications 12: 1406-1417.
 

 

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See more publications of ITES Faculty

 

 

  Biol. 6995 - Biogeochemistry

During the last two decades, tropical environmental issues have attracted a great deal of scientific and public attention.  These issues include, for example, tropical deforestation linked with global warming, and land use changes associated with sustainability of plant primary productivity and maintenance of tropical biodiversity.  This course will introduce some of the fundamental biogeochemical principles which are used in studies of tropical ecological issues.  A strong theme of this class will examine how natural biogeochemical processes operate and how human activities can alter these processes.  The subject has been divided roughly into two parts: natural biogeochemical processes, and interactions between natural and anthropogenic processes.  In the first part, we will discuss biogeochemical processes occurring in atmosphere, land, and in ocean.  In the second part, we will use all that covered thus far to understand the anthropogenic effects on the cycling of carbon, nitrogen, phosphorus, cations, and sulfur and the implications to a variety of environmental issues.  Thus the course will analyze biogeochemical cycles from several different points of view, emphasizing an integrated approach.

Biol. 3111 - General Ecology

Teach the fundamentals of ecology to non-ecology majors. The course covers all aspects of ecology, from autoecology, developmental and behavioral ecology, through population and community ecology, ecosystems, biogeography and global change.

 

 

  Matthew Warren
    mwarren1@lternet.edu

 

Project:
Nutrient retaintion in tropical forests
 


  Sara Raquel
    namanereis@yahoo.com

 

Projects:

Soil doc in tropical pastures and forests

 

 

  Office: 
Facundo Bueso annex-208 (XXX)

UPR extensión number:
787 764-0000 x 2868

Fax number:
787-772-1481